Large larvae, up to 15 mm long.
Antenna with ring organ in basal 1/3 of basal segment. Blade longer than flagellum.
Labrum with SI with apical teeth; other S setae normal. Labral lamellae broad, pectinate. Chaetae serrated; chaeta media conspicuous. With about 3-4 spinulae. Pecten epipharyngis consisting of 3 scale-like spines. Chaetulae laterales smooth; chaetulae basales well-developed, bifid at apex. Premandible with 2 apical teeth and weak brush.
Mandible with apical tooth longer than combined width of 4 inner teeth. Seta interna with serrate branches.
Mentum with 12-14 teeth or 16-18 teeth including the 2 accessory teeth on each side of 1st lateral teeth; 2 median teeth low; 1st lateral tooth high, trifid; 4th and 6th (when present) lower than 5th. Ventromental plate broad and long, Chironomini-like, numerous strong setae in beard.
Maxilla with anterior lacinial chaeta not conspicuous. Lamellae of galea scale-like, triangular, smooth. Cardinal beard present.
Body without conspicuous body setae.
Larvae in springs, streams, rivers, pools, ponds, and the littoral zone of lakes. Holarctic. Four of five species described as larvae.