Notocladius Harrison

Description

Introduction

Very small larvae, up to 2 mm long.

Antenna

Antenna 4 segmented; segments 3 and 4 subequal, combined about 50% length of 2nd. blade extending to apex of second segment. Lauterborn organs small but distinct, opposite on apex of 2nd segment.

Labrum

Labrum with SI bifid, other setae simple; quite strong and almost lanceolate, SII and SII arising from small but disticnt tubercles. SIV undescribed. 1 pair bifid chaetae. Pecten epipharyngis consisting of 3 rounded scales; three pairs of chaetulae laterales each broad and with 'ciliary margin' (Harrison, 1997), chaetulae basales apparently absent. Premandible broad, with 1 large broad apical tooth, small outer tooth, and distinct, sinuous brush.running along outer margin from at least midpoint to extend slightly beyond apex.

Mandible

Mandible with apical tooth subequal to first of 4 inner teeth. Seta subdentalis weak. Seta interna with 5-6 weakly divided, almost hyaline, branches.

Mentum

Mentum with 3 median teeth and 5 pairs of lateral teeth. Ventromental plate poorly developed anteriorly, abruptly making 90 degree angle to direct laterally, without beard beneath. Setae submenti stout.

Maxilla

Maxilla with small sensillae on palp, larger sensillae on galea. Galear surface with several lamellae Pecten galearis and appendix absent.

Body

Body with meso- and metathoracic segments fused; only 2 distinctive thoracic segments present, although long 'neck' provides appearance of a first segment with anterior parapods located on swollen 'second'. Anterior and posterior parapods separate, squat, and bearing simple claws. Procercus very short, small and bearing apically 4 short setae. Anal tubules rounded and projecting a little further than posterior parapods. A stout, simple, short seta arises from ventral basal half of posterior parapods.

Notes

Taxonomic Notes

The larvae are distinctive by size (small) and somewhat 'comma-shaped' appearance. The thorax is swollen such that it is near half the body length, the neck is quite well developed, and the short paired anterior parapods are situated quite posteriorly, actually on the well developed prothorax, anterior to a fused meso- and meta-thorax. In these features it is clear that Notocladius belongs with Thienemanniella and Corynoneura.

Ecological Notes

Known from one species, N. capicola, from the Cape area of South Africa. Larvae live on rock faces in fast running waters, even waterfalls, in clear, clean mountain and submontane streams (Harrison, 1997) from the Cape fold to the Cederberg Mountains.

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